Rotary International reaches 1.4 million members across 200 countries and territories, with a public health and water legacy that helped drive polio to the edge of eradication. The 7 Seas Foundation is building parallel infrastructure for ocean literacy and marine conservation. This fireside explores how legacy civil society networks are being retooled for the nature agenda — the operational lessons from water, sanitation, and disease programs, and how they transfer to ocean and watershed resilience at continental scale.